These variations have been explained through the differences in density, surface area, and the size of plastic litter ( Chubarenko et al., 2016 Kowalski et al., 2016 Schwarz et al., 2019). Variation in quantity and composition was observed throughout the different environmental compartments: polyethylene (PE) and polypropylene (PP) were mostly observed in epipelagic waters, whereas polyamide and polyester dominated in sediments. Vast accumulation zones have been identified in the five subtropical oceanic gyres ( Van Sebille et al., 2015), but also in the Mediterranean Sea that has been proposed as the sixth great accumulation zone for marine litter ( Cózar et al., 2015). In this area, the longevity of the plastics is estimated to be hundreds to thousands of years ( Barnes et al., 2009 Lusher et al., 2015 Kane et al., 2020). Plastic pollution has become a global environmental problem affecting all parts of oceans worldwide, including the most remote areas such as deep seafloor or polar regions. Our results showed that the plastic size and shape had relatively low influence on the plastisphere abundance, diversity, and activity, as compared to the plastic composition or the presence of a phytoplankton bloom. The diatom bloom event had a great impact on the plastisphere of all materials, resulting in a drastic change in diversity and activity. A mature biofilm developed within 30 days on all material types, with higher bacterial abundance on the plastics compared to glass, and distinct bacterial assemblages were detected on each material type. Bacterial communities associated with the material of 100 μm size fraction showed the highest activity compared to all other material sizes. However, plastic size and shape had an effect on bacterial activity ( 3H leucine incorporation). Results of bacterial abundance (confocal microscopy) and diversity (16S rRNA Illumina sequencing) indicated that the three classical biofilm colonization phases (primo-colonization after 3 days growing phase after 10 days maturation phase after 30 days) were not influenced by the size and the shape of the materials, even when a diatom bloom ( Pseudo-nitzschia sp.) occurred after the first month of incubation. Polyethylene (PE) and polylactide acid (PLA) together with glass controls in the forms of meso-debris (18 mm diameter) and large-microplastics (LMP 3 mm diameter), as well as small-microplastics (SMP) of 100 μm diameter with spherical or irregular shapes were immerged in seawater during 2 months. Here, we aimed to evaluate the relative influence of the plastic size, shape, chemical composition, and environmental changes such as a phytoplankton bloom in shaping the plastisphere abundance, diversity and activity. The thin film of life that inhabits all plastics in the oceans, so-called “plastisphere,” has multiple effects on the fate and impacts of plastic in the marine environment. 6SAS Observatoire Océanologique de Banyuls-sur-Mer, Banyuls-sur-Mer, France.5IMRCP, Université de Toulouse, CNRS, Toulouse, France.4Institut de Recherche Dupuy de Lôme (IRDL), Université Bretagne Sud, UMR CNRS 6027, Lorient, France.3Laboratoire Charles Coulomb (L2C), UMR 5221 CNRS-UM, Place Eugène Bataillon, Montpellier, France.
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